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The Mathematical understanding has been studied before, but no research has a focus on mathematical understanding and adversity quotient (AQ) of the pre-service mathematics teacher. This study is experimental research that aims to know about the influence of AQ of pre-service mathematics teacher toward the achievement of mathematical understanding ability. The population of this study is the pre-service mathematics teacher in Cimahi City, West Java, Indonesia; while the sample is 55 pre-service mathematics teachers selected purposively. The results show that: (1) AQ gives positive influence to the development of mathematical understanding ability of pre-service mathematics teacher with the influence of 57.3%, while the rest of it (42.7%) is influenced by other factors outside AQ; (2) there are differences in achievement in mathematical understanding ability of pre-service mathematics teacher based on AQ level (climber, camper, quitter).